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Density-size dynamics of trees simulated by a one-sided competition multi-species model of rain forest stands

, : Density-size dynamics of trees simulated by a one-sided competition multi-species model of rain forest stands. Annals of Botany (London) 70(5): 451-460

Stand development processes from various initial densities were simulated employing a one-sided competition model of the size structure dynamics of trees in which the size growth rate and the probability of survival of a tree are linearly depressed by the cumulative basal area of trees in larger size classes. All model functions and parameters were derived from permanent plot censuses in a warm-temperate rain forest in southern Japan. The simulated tree density-mean tree size trajectories using species-averaged parameters were similar to those previoulsy reported for even-aged monocultures. An upper boundary of the density-size trajectories or the self-thinning line did not change when the recruitment of seedlings was taken into account. The observed variance of growth rate at a given size hardly affected the results of simulation. A simulation of multi-species system composed of the three most abundant species showed the same density-size dynamics in terms of total yield as a species-averaged system, while not in terms of each species cohort. It is concluded that the established rules of competition-density effect and self-thinning in higher plants are due to one-sided competition among co-occurring plant individuals of similar life forms irrespective of species.

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