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Systematics of the terpides lineage from the neotropics definition of the terpides lineage methods and revision of fittkaulus savage and peters insecta ephemeroptera leptophlebiidae

, : Systematics of the terpides lineage from the neotropics definition of the terpides lineage methods and revision of fittkaulus savage and peters insecta ephemeroptera leptophlebiidae. Spixiana 9(3): 255-270

The monophyletic Terpides lineage is established, defined, and distinguished from all other genera of Leptophlebiidae by characters in the imaginal and nymphal stages. This lineage presently includes Terpides Demoulin, Fittkaulus Savage and Peters, and an undescribed genus. The Terpides lineage stems from the base of the Atalophlebiinae very near the basal furcation within the Leptophlebiidae between the subfamilies Atalophlebiinae and Leptophlebiinae. Members of the Terpides lineage were isolated on the ancient Guiana and Brazilian Shields of South America during the late Cretaceous and early Tertiary and dispersed northward when the isolation of South America ended with the development of the lower Central American isthmus in Pliocene-Pleistocene times. The genus Fittkaulus is revised and two new species are described [F. cuiabae (Brazil) and F. cururuensis (Brazil)]. Fittkaulus is currently known from the eastern Amazon Basin, from river basins draining north from the Eastern Guiana Highlands, and from the northern portion of the Rio Paraguay Basin in Brazil.

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