The pattern of distribution of dry matter and P was followed in individual plants harvested from a field crop of spring wheat throughout its development. There was a continued P uptake with time and almost half of the total P was accumulated in the post-anthesis period of growth. The various component parts of the main shoot were followed in detail and both individual leaves and the stem showed substantial net losses of P with time well before they reached their dry weight maxima. The ear was the major sink for P and it was estimated that approximately 1/3 of its P content was supplied by translocation. The main shoot stem and leaves also showed a significant decline in dry weight during the final stages of grain development. Half-ear removal at anthesis increased both grain set and the growth of the grain of the upper florets of the remaining spikelets. Defoliation had little effect on grain yield but resulted in a reduction in the P content of the grain.